The "heat increment" associated with feeding and digestion was estimated as 15 to 17% of the oxygen uptake by M. edulis during all stages of recovery. M. dedulis adapted to the intertidal regime by reducing its time-averaged aerial and aquatic rates of ammonia excretion. In contrast, C. edule maintained a high aerial and aquatic rate of ammonia excretion. The clearance rates of M. edulisrecovered rapidly (0.5 to 1.5 h) following reimmersion, whereas those of C. edule recovered more slowly, particularly for the subtidal individuals. Intertidally acclimatized individuals of M. edulis and C. edule had more energy available for growth (scope for growth) integrated over a 12 h period and higher growth efficiencies than subtidal individuals subjected to acute air exposure and intermittent feeding regimes.
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